The dragons and their dedicated band of keepers have at last found the lost city of Kelsingra. The magical creatures have learned to use their wings and are growing into their regal inheritance. Their humans, too, have begun transforming into beautiful Elderlings. But while the humans have scoured the empty streets and enormous buildings of Kelsingra, they cannot find the mythical silver wells the dragons need to stay healthy and survive. And time is of the essence, for the legendary Tintaglia, long feared dead, has returned, wounded in a battle. She is weakening and only the hidden silver can revive her. If Tintaglia dies, so too will the ancient memories she carries - a devastating loss that will ensure the dragons' extinction.